Our run defense AI will now have the ability to read repeated plays ran by the offense and react in a more realistic manner. Defenses will start to key in on repetitive offensive run plays. If a team is running the same play out of the same formation, that is a tell to the defense. Once the defense keys in on this, defenders will play their run-fit and start hitting their gap as soon as the ball is snapped. This will not apply on a play-action run.
How quickly defenders begin to adjust will be dictated by the skill level the user is playing on. On Rookie difficulty, it may take the defense more time to catch on and predict what play is coming compared to our All-Madden difficulty where the defense may only need 1 play before they make effective adjustments.
Your players and ratings are still a huge factor here. An example of this would be that a Defensive Lineman with lower blockshed and strength ratings will still be to be outmatched by an Offensive Lineman with higher blocking ratings, resulting in still having a better chance at protecting the ball carrier in the run game.
The difference here is that the defense will not make the same mistakes as the offense runs what they have seen consistently. All of this will result with a more authentic experience as the defense responds just like how their real life counterparts would.